Conclusions

Could these two genetic defects, absence of chlorophyll and absence of anthocyanins, be linked? We see one plant with both pigments absent growing next to another with one absent. Either there is a relationship or it is coincidence.

If there is a link between the two supposed genes involved they would be located on the same chromosome; the closer they are the more linkage. Remember that cross-over of chromosomal material during meiosis can segregate genes. The closer they are the more likely that they are inherited together. However, if this was the case then we would not see just anthocyanin deficiency as shown by var. monotropoides. The two abnormalities must be inherited independently.
There must be, in the parent population of both plants, those with affected genes for both pigment deficiencies, and the random inheritance created by the randomness of cross-pollination. Some of the other Broad-leaved Helleborines at the site, and perhaps locally, are therefore likely to be heterozygotes (carriers) for either abnormality and it is possible more examples of such plants may appear, including a E. h. var. chlorantha.

Should this plant be termed Epipactis helleborine var. chlorantha x var. monotropoides?